Contents of Draft Document on Deer Culling, Part 1 (Background) 1.1 How many deer are there in the UK? 1.2 What is the effect of an overpopulation of deer? Do we need to keep numbers down? 1.3 Can deer be kept out of the areas where they can cause damage? 1.4 So if there are too many deer, can we improve woodland regeneration by reducing their numbers? 1.5 So how are deer culled? --------------- 1. Background 1.1 How many deer are there in the UK? --------------------------- There are more than a million deer in mainland UK. This includes 500,000 Roe Deer, 360,000 Red Deer, 100,000 Fallow Deer and 40,000 Muntjac - from the 1995 JNCC Review of British Mammals). The total number of deer has risen dramatically in recent years. Estimates were made in the early 1970s of 200,000 Roe Deer, 190,000 Red Deer, 50,000 Fallow Deer and 5,000 Muntjac, though it is acknowledged that these were less accurate than more recent figures. Deer have no natural predators in the UK, save man, and uncontrolled their population will continue to rise. 1.2 What is the effect of an overpopulation of deer? Do we need to keep numbers down? ------------------------ The detrimental effects of an overpopulation of deer on our woodlands are well documented. Any ecologist will tell you the value of preventing any species in a woodland from getting out of control. If any of the herbivores get too numerous, the damage done can be tremendous. Some reference to the literature published in this field shows the necessity of maintaining an appropriate population: Buckley GP, Howell R, Watt TA, et al. Vegetation succession following ride edge management in lowland plantations and woods .1. The influence of site factors and management practices BIOL CONSERV 82: (3) 289-304 DEC 1997 This paper gives a study of woodland regeneration. "Deer grazing profoundly affected vegetation composition and structure, greatly reducing tree and shrub regeneration. " Mayle BA Progress in predictive management of deer populations in British woodlands FOREST ECOL MANAG 88: (1-2) 187-198 NOV 1 1996 This is a very important study on deer control. Interestingly, there's more emphasis on the level of population control than on the necessity for population control, the case for the necessity of control being assumed to be beyond dispute. Kirby KJ, Thomas RC, Dawkins HC Monitoring of changes in tree and shrub layers in Wytham woods (Oxfordshire), 1974-1991 FORESTRY 69: (4) 319-334 1996 Looks at a wood that has basically thinned a little in the time period, exploring the reasons why. One of the factors was grazing by deer. "The shrub cover has also declined greatly, probably because of increased deer browsing." Patel A, Rapport DJ Assessing the impacts of deer browsing, prescribed burns, visitor use, and trails on an oak-pine forest: Pinery Provincial Park, Ontario, Canada NAT AREA J 20: (3) 250-260 JUL 2000 A recent study of the effect of high deer population density on woodland. "Higher deer densities led to significant declines in species richness, stem density, cover, and median seedling height." Scott D, Welch D, Thurlow M, et al. Regeneration of Pinus sylvestris in a natural pinewood in NE Scotland following reduction in grazing by Cervus elaphus FOREST ECOL MANAG 130: (1-3) 199-211 MAY 1 2000 Looks at whether the initial regeneration of a gap in a forest is affected by a number of factors. Found that the presence of red deer, grazing the area, could give saplings a chance of germinating, but did not demonstrate that those saplings had any increased chance of surviving to adulthood, and the statistical increase in germination is very low. Jorritsma ITM, van Hees AFM, Mohren GMJ Forest development in relation to ungulate grazing: a modelling approach FOREST ECOL MANAG 120: (1-3) 23-34 JUL 12 1999 "The results presented in this paper concern the development of a pine forest (Pinus sylvestris L.) in the Netherlands under various grazing pressures over a period of 100 years. They show that even low densities of ungulates can have significant impacts on the regeneration and thereby on forest development. " Radeloff VC, Pidgeon AM, Hostert P Habitat and population modelling of roe deer using an interactive geographic information system ECOL MODEL 114: (2-3) 287-304 JAN 1 1999 Discusses the level to which culling is necessary. "We describe a model to determine deer population densities compatible with forest management goals, and to assess harvest rates necessary to maintain desired deer densities. " Impact of bark stripping by sika deer, Cervus nippon, on subalpine coniferous forests in central Japan Yokoyama N, Maeji I, Ueda T, Ando M, Shibata E FOREST ECOLOGY AND MANAGEMENT 140: (2-3) 93-99 JAN 15 2001 Demonstrates that a major killer of trees was sika deer. By stripping too much bark, trees died, which led to a long term change in the species of plants represented in those forests and an overall loss of woodlands. Online information sources also comment on the effects of having more large herbivores than can be sustained in a habitat. The Scottish Wildlife Trust for example have published a statement on the necessity of their culling policy. (http://www.swt.org.uk/Publications/Policies/reddeerpol.htm) "Red deer may also cause serious damage to blanket bogs, by wallowing in pools and in association with grazing sheep, are responsible for many arctic-alpine plants (e.g.: Alpine Sow Thistle) being restricted to inaccessible rocky outcrops and ledges. Scotland's Red Deer population is forever increasing (from around 216,000 in the early 1960s to over 300,000 in 1986), so it is likely that the environmental problems associated with them will also continue to increase in size. A solution must be found soon." In the year 2000, 71 000 deer were culled in Scotland. Despite this, the population continues to rise (http://213.235.21.10/LifeCountry/Pursuits/shoot_news6.htm). Although many animal rights activists undoubtedly find the culling of deer to maintain sustainable population density quite distasteful, organisations such as the League Against Cruel Sports (http://www.league.uk.com/) accept the necessity of this practice. The largest, and most famous population of deer in England is within the Exmoor National Park. With the possible demise of hunting of deer with hounds in that area, the Burns enquiry concluded that the development of an alternative culling strategy to maintain numbers at a sustainable level is of critical importance (quoted at: http://www.parliament.the-stationery-office.co.uk/pa/cm199900/cmhansrd/vo000 707/debtext/00707-05.htm) 1.3 Can deer be kept out of the areas where they can cause damage? ----------------------- In some instances, fences can be used to keep deer out. This is especially useful when new saplings have been planted. Unfortunately, this doesn't provide a complete answer. The choice of how to protect your saplings comes down to either individual tree protection or fencing of a whole area. Fencing can be problematic, especially in Scotland, for a number of reasons. One of the emblems of the Scottish countryside is the capercaillie. This bird is unfortunately endangered, its numbers severely restricted having halved in the last ten years to no more than 1000 individuals. One of the reasons for this is the use of deer fencing within and around areas of woodland. The birds, adults and young, fly low between the trees, hit the fences and die. This has been demonstrated in the following research papers: Capercaillie Tetrao urogallus in Scotland - demography of a declining population Moss_R, Picozzi_N, Summers_RW, Baines_D IBIS, 2000, Vol.142, No.2, pp.259-267 And Assessment of bird collisions with deer fences in Scottish forests Baines_D, Summers_RW JOURNAL OF APPLIED ECOLOGY, 1997, Vol.34, No.4, pp.941-948 Quotes from this report: "This study adds weight to previous findings that fences are a frequent cause of mortality in capercaillie", and "a policy of deer culling to achieve tree regeneration without deer fencing would be desirable wherever practicable and especially within the range and main native pinewood habitat of capercaillie". One RSPB document (http://www.ierm.ed.ac.uk/restore/proc8.htm) states : "We now know that the effect of deer fences on woodland grouse has been very severe. Fences are probably one of the main reason why capercaillie have declined from around 20,000 birds in the 1970's to about 2,200 birds at the present time. At Abernethy, as a result of fence removal and improvements in habitat, there has been a 300% increase in the number of lekking males" Quoting again from the Scottish Wildlife Trust (http://www.swt.org.uk/Publications/Policies/reddeerpol.htm) "While deer fencing can exclude deer from specific areas, and can be used to promote (for example), woodland regeneration it has many drawbacks. It is expensive to erect, concentrates grazing deer at a higher density in areas beyond the fence, it can cause fatalities amongst birds such as capercaillie (when they fly into fences), and unless it is very well planned it can cause access problems in areas used for outdoor recreation." The above article continues with a discussion of the problems with deer climbing over the fences in snowy conditions and becoming trapped. The alternative to this is individual tree protection. While this can be done to protect saplings in new plantations, it is extraordinarily expensive. A quote from a private contractor, with the comparative cost of protected and unprotected planting, is shown below Comparative typical planting costs of 40 - 60 cm. native tree saplings. All figures at per 1000 rate. Costs may vary according to scale, conditions, stocking density and management company. Bare planting: Tree - 40 - 60 cm. 15p +VAT + delivery Labour 9p Profit @ 20% 5p +VAT 29p Optimum planting density 2250 per ha. @ 29p = £652:50 per ha. Planting in standard 1.2 m 'tubes' (effective against roe deer) Tree - 40 - 60 cm. 15p +VAT + delivery Standard tube 67p +VAT + delivery 4'6" planting stake 31p +VAT + delivery Labour 45p +VAT Removal of tube and stake at 5-8 years 11p +VAT Transport and disposal of tubes. 3p +VAT Profit @ 20% 34p +VAT £2:06p 2250 per ha @ £2:06p = £4,635 per ha Plus environmental costs of tube production and landfill. Labour costs for replacing any failed trees in years 1 and 2 are also much higher than for bare planted trees because each tube has to be checked individually, removed and re-attached during replacement planting. Thus, planting trees with individual protection is more than seven times more expensive. While preventing the deer from eating the saplings, this technique does nothing to reduce deer numbers. The result can be an overgrazed forest floor, with little species diversity, and a lack of any natural regeneration of trees. 1.4 So if there are too many deer, can we improve woodland regeneration by reducing their numbers? -------------------------------------- Deer do not have to be completely eliminated to allow woodlands to regenerate. In the RSPB reserve at Abernathy (http://www.ierm.ed.ac.uk/restore/proc8.htm) a reduction of deer numbers from 12 per hectare to less than four had a radical effect on woodland regeneration. "In one area of Abernethy, a scatter of pine trees were left after a big felling during the 1st World War. On this site there has been no regeneration for the last 100 years. As a result of our deer management, around some of these old trees there are little pockets of regeneration starting to appear." The recover of important shrub communities, with species such as dwarf birch and juniper, has also been recorded at this site. But it's not just the plant community that prospers when deer numbers are reduced to a sustainable level. As mentioned above, at Abernathy the numbers of important bird species have increased massively as a result of maintaining a sustainable number of deer. Further information on this project can be found in this reference: Beaumont, D., Dugan, D., Evans, G. and Taylor, S. (1995): Deer management and tree regeneration in the RSPB reserve at Abernethy Forest. Scottish Forestry, 49(3), 155-161 1.5 So how are deer culled? ---------------------------- The Association of Deer Management Groups has strict guidelines for how this is achieved (http://www.deer-management.co.uk/). "This is undertaken by professional stalkers who have unmatched experience in working with wildlife in Scotland, and a close knowledge of deer in particular. The culling of deer involves the selection of older or unhealthy animals which are shot humanely with a high velocity rifle. Culling policy is agreed by Deer Management Groups and allocated among their members on the basis of a regular coordinated count by the Group, or by the Deer Commission for Scotland" The welfare implications of killing red deer by this method were explored in depth in this work. Welfare implications of culling red deer (Cervus elaphus) Bradshaw EL, Bateson P ANIMAL WELFARE 9: (1) 3-24 FEB 2000 This study found that almost 90% of deer were killed with a single shot, and that only 7 percent took between 2 and 15 minutes to die. Only about 2% of animals escaped wounded. Despite this high success rate, the study concluded that there is still scope to improve on this.
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